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論文


2024

  •         Ayaka Saito, Chihiro Kinoshita, Kino Sakai, Katsufumi Sato, Kentaro Q. Sakamoto. Heart rate reduction during voluntary deep diving in free-ranging loggerhead sea turtles. Journal of Experimental Biology 227: jeb246334 (2024).
  •         Daiki Uematsu, Takuto Kishimoto, Kazuki Harada, Shinichi Watanabe, Katsufumi Sato, Hidetoshi Takahashi. Nest box for monitoring the mass and heart rate changes of a growing seabird chick. IEEJ Transactions on Sensors and Micromachines 144: 1–12 (2024).


2023

  •         Makoto A. Yoshida, Mabuchi K. Using a drifting GPS-Argos satellite buoy as a method for detecting acoustic-tagged fish offshore in an ancient lake. Animal Biotelemetry 11:30 (2023).
  •         Bethany Clark, Ana Carneiro, Elizabeth Pearmain, Marie-Morgane Rouyer, Thomas Clay, Win Cowger, Richard Phillips, Andrea Manica, Carolina Hazin, Marcus Eriksen, Jacob González-Solís, Josh Adams, Yuri Albores-Barajas, Joanna Alfaro-Shigueto, Maria Alho, Deusa Araujo, José Manuel Arcos, John Arnould, Nadito Barbosa, Christophe Barbraud, Annalea Beard, Jessie Beck, Elizabeth Bell, Della Bennet, Maud Berlincourt, Manuel Biscoito, Oskar Bjørnstad, Mark Bolton, Katherine Booth Jones, John Borg, Karen Bourgeois, Vincent Bretagnolle, Joël Bried, James Briskie, Michael Brooke, Katherine Brownlie, Leandro Bugoni, Licia Calabrese, Letizia Campioni, Mark Carey, Ryan Carle, Nicholas Carlile, Ana Rita Carreiro, Paulo Catry, Teresa Catry, Jacopo Cecere, Filipe Ceia, Yves Cherel, Chang-Yong Choi, Marco Cianchetti-Benedetti, Rohan Clarke, Jaimie Cleeland, Valentina Colodro, Bradley Congdon, Jóhannis Danielsen, Federico De Pascalis, Zoe Deakin, Nina Dehnhard, Giacomo Dell'Omo, Karine Delord, Sébastien Descamps, Ben Dilley, Herculano Dinis, Jérôme Dubos, Brendon Dunphy, Louise Emmerson, Ana Fagundes, Annette Fayet, Jonathan Felis, Johannes Fischer, Amanda Freeman, Aymeric Fromant, Giorgia Gaibani, David García, Carina Gjerdrum, Ivandra Soeli Gomes, Manuela Forero, José Pedro Granadeiro, W Grecian, David Grémillet, Tim Guilford, Gunnar Hallgrimsson, Luke Halpin, Erpur Hansen, April Hedd, Morten Helberg, Halfdan Helgason, Leeann Henry, Hannah Hereward, Marcos Hernández-Montero, Mark Hindell, Peter Hodum, Simona Imperio, Audrey Jaeger, Mark Jessopp, Patrick Jodice, Carl Jones, Christopher Jones, Jón Einar Jónsson, Adam Kane, Sven Kapelj, Yuna Kim, Holly Kirk, Yann Kolbeinsson, Philipp Kraemer, Lucas Krüger, Paulo Lago, Todd Landers, Jennifer Lavers, Matthieu Le Corre, Andreia Leal, Maite Louzao, Jeremy Madeiros, Maria Magalhães, Mark Mallory, Juan Masello, Bruno Massa, Sakiko Matsumoto, Fiona McDuie, Laura McFarlane Tranquilla, Fernando Medrano, Benjamin Metzger, Teresa Militão, William Montevecchi, Rosalinda Montone, Leia Navarro-Herrero, Verónica Neves, David Nicholls, Malcolm Nicoll, Ken Norris, Steffen Oppel, Daniel ORO, Ellie Owen, Oliver Padget, Vitor Paiva, David Pala, Jorge Pereira, Clara Péron, Maria Petry, Admilton Pina, Ariete Teresa Pina, Patrick Pinet, Pierre Pistorius, Ingrid Pollet, Benjamin Porter, Timothée Poupart, Christopher Powell, Carolina Proaño, Júlia Pujol-Casado, Petra Quillfeldt, John Quinn, Andre Raine, Helen Raine, Iván Ramírez, Jaime Ramos, Raül Ramos, Andreas Ravache, Matt Rayner, Timothy Reid, Gregory Robertson, Gerard Rocamora, Dominic Rollinson, Robert Ronconi, Andreu Rotger, Diego Rubolini, Kevin Ruhomaun, Asunción Ruiz, James Russell, Peter Ryan, Sarah Saldanha, Ana Sanz-Aguilar, Mariona Sardà Serra, Yvan Satgé, Katsufumi Sato, Wiebke Schäfer, Stefan Schoombie, Scott Shaffer, Nirmal Shah, Akiko Shoji, Dave Shutler, Ingvar Atli Sigurðsson, Mónica Silva, Alison Small, Cecilia Soldatini, Hallvard Strøm, Christopher Surman, Akinori Takahashi, Rabindra Vikash Tatayah, Graeme Taylor, Robert Thomas, David Thompson, Paul Thompson, Thorkell Thórarinsson, Diego Vicente-Sastre, Eric Vidal, Ewan Wakefield, Susan Waugh, Henri Weimerskirch, Heiko Wittmer, Takashi Yamamoto, Ken Yoda, Carlos Zavalaga, Francis Zino, and Maria Dias. Global assessment of marine plastic exposure risk for oceanic birds. Nature Commumications 14: 3665 (2023).
  •         Miho Sakao, Tomoko Hamabata, Katsufumi Sato, Shinichi Watanabe, Ken Yoda, Kozue Shiomi. Absence of genetic structure among Streaked Shearwaters Calonectris leucomelas breeding in Japan, despite limited dispersal events. Ornithological Science 22: 111-121 (2023).
  •         Ryotaro Manabe, Takatoshi Higuchi, Shun Watanabe, Fadly Y. Tantu, Hagi Y. Sugeha, Hiroki Kaneko, Michael J. Miller, Seishi Hagihara, Tatsuki Yoshinaga, Augy Syahailatua, Sam Wouthuyzen, Triyanto, Kawilarang W. A. Masengi, Katsufumi Sato, Jun Aoyama, Katsumi Tsukamoto. Migration behavior of Anguilla celebesensis silver eels within their Tomini Bay spawning area. Zoological Studies 62: 46 (2023).
  •         Leo Uesaka, Yusuke Goto, Masaru Naruoka, Henri Weimerskirch, Katsufumi Sato, Kentaro Q. Sakamoto. Wandering albatrosses exert high take-off effort only when both wind and waves are gentle. eLife 12:RP87016 (2023).
  •         Takashi Iwata, Kagari Aoki, Patrick J. O. Miller, Martin Biuw, Michael J. Williamson, Katsufumi Sato. Non-lunge feeding behaviour of humpback whales associated with fishing boats in Norway. Ethology 2023: 1-6 (2023).
  •         Masao Amano, Kagari Aoki, Hayao Kobayashi, Shingo Minamikawa, Katsufumi Sato, Tsunemi Kubodera. Stereotypical diel movement and dive pattern of male sperm whales in a submarine canyon revealed by land-based and bio-logging surveys. Frontiers in Marine Science 10: 1150308 (2023).
  •         Chihiro Kanno, Kentaro Q. Sakamoto, Sung-Sik Kang, Yojiro Yanagawa, Seiji Katagiri, Masashi Nagano. The difference in subpopulation structures in sex-sorted and non-sorted semen by discriminant analysis of bull sperm motility data collected by a computer-assisted sperm analysis system. Andrologia. 2023: 9944344 (2023).


2022

  •         Takuya Fukuoka, Hiroyuki Suganuma, Satomi Kondo, Katsufumi Sato. Long dive capacity of olive ridley turtles (Lepidochelys olivacea) at high water temperature during the post-nesting foraging period in the Arafura Sea. Journal of Experimental Marine Biology and Ecology546: 151649 (2022).
  •         Leo Uesaka, Yusuke Goto, Yoshinari Yonehara, Kosei Komatsu, Masaru Naruoka, Henri Weimerskirch, Katsufumi Sato, Kentaro Q. Sakamoto. Ocean wave observation utilizing motion records of seabirds. Progress in Oceanography 200: 102713 (2022).
  •         Shigenori Nobata, Katsufumi Sato, Shouji Houki, Motohiro Ito, Yoshinori Aoki, Takashi Kitagawa and Susumu Hyodo. Straightforward upriver migration to spawning sites in chum salmon Oncorhynchus keta homed to the coastal short river of the Sanriku region. Journal of Fish Biology 2022.1-10 (2022).
  •         Chihiro Kinoshita, Ayaka Saito, Megumi Kawai, Katsufumi Sato, Kentaro Q. Sakamoto. A non-invasive heart rate measurement method is improved by placing the electrodes on the ventral side rather than the dorsal in loggerhead turtles. Frontiers in Physiology 13:811947 (2022).
  •         Takuya Fukuoka, Fumiki Sakane, Chihiro Kinoshita, Katsufumi Sato, Kaoruko Mizukawa, Hideshige Takada. Covid-19-derived plastic debris contaminating marine ecosystem: Alert from a sea turtle. Marine Pollution Bulletin 175: 113389 (2022).


2021

  •         Kentaro Q. Sakamoto, et al. A non-invasive system to measure heart rate in hard-shelled sea turtles: Potential for field applications. Philosophical Transaction 376: 20200222 (2021).
  •         Clive R. McMahon, Fabien Roquet, Sophie Baudel, Mathieu Belbeoch, Sophie Bestley, Clint Blight, Lars Boehme, Fiona Carse, Daniel P. Costa, Mike Fedak, Christophe Guinet, Robert Harcourt, Emma Heslop, Mark A Hindell5, Xavier Hoenner, Kim Holland, Melinda Holland, Fabrice R. A Jaine, Tiphaine Jeanniard du Dot, Ian Jonsen, Theresa R. Keates, Kit M. Kovacs, Sara Labrousse, Philip Lovel, Christian Lydersen, David March, Matthew Mazloff, Megan K. McKinzie, Mônica M. C. Muelbert, Kevin O’ Brien, Lachlan Phillips, Esther Portela, Jonathan Pye, Steve Rintoul, Katsufumi Sato, Ana M. M. Sequeira, Samantha Simmons, Vardis M. Tsontos, Victor Turpin, , Esmee van Wijk, Danny Vo, Mia Wege, Fred Whoriskey, Kenady Wilson, Bill Woodward. Animal borne ocean sensors -AniBOS- An essential conponent of the global ocean observing system. Frontiers in Marine Science 8: 751840. (2021).
  •         Kagari Aoki, Saana Isojunno, Charlotte Bellot, Takashi Iwata, Joanna Kershaw, Yu Akiyama, Lucía Martina Martín López, Christian Ramp, Martin Biuw, René Swift, Paul Wensveen, Patrick Pomeroy, Tomoko Narazaki, Ailsa Hall, Katsufumi Sato, Patrick J. O. Miller. Aerial photogrammetry and tag-derived tissue density reveal patterns of lipid-store body condition of humpback whales on their feeding grounds. Proceedings of Royal Society B 288: 20202307 (2021).
  •         Chihiro Kinoshita, Takuya Fukuoka, Tomoko Narazaki, Yasuaki Niizuma and Katsufumi Sato. Analysis of why sea turtles swim slowly: a metabolic and mechanical approach. Journal of Experimental Biology 224: jeb236216 (2021).
  •         Hiromi Kudo, Kei Uchida, Hideaki Nishizawa, Katsufumi Sato. Boldness-exploration behavioral syndrome in wild sub-adult green sea turtles caught at Oita, Japan. Applied Animal Behaviour Science 236: 105216 (2021).
  •         Tomoko Narazaki, Itsumi Nakamura, Kagari Aoki, Takashi Iwata, Kozue Shiomi, Paolo Luschi, Hiroyuki Suganuma, Carl G. Meyer, Rui Matsumoto, Charles A. Bost, Yves Handrich, Masao Amano, Ryosuke Okamoto, Kyoichi Mori, Stéphane Ciccione, Jérôme Bourjea, Katsufumi Sato. Similar circling movements observed across marine megafauna taxa. iScience 24: 102221 (2021).
  •         Takashi Iwata, Martin Biuw, Kagari Aoki, Patrick James O’Malley Miller, Katsufumi Sato. Using an omnidirectional video logger to observe the underwater life of marine animals: humpback whale resting behaviour. Behavioural Processes 186: 104369 (2021).
  •         Masaru Naruoka, Yusuke Goto, Henri Weimerskirch, Takashi Mukai, Taichi Sakamoto, Kentaro Q. Sakamoto, Katsufumi Sato. Application of inertial and GNSS integrated navigation to seabird biologging. Journal of Robotics and Mechatronics 33: 526-536 (2021).
  •         Hidetoshi Takahashi, Masaru Naruoka, Yoshinobu Inada, Katsufumi Sato. Seabird biologging system with compact waterproof airflow sensor. Journal of Robotics and Mechatronics 466-476 (2021).
  •         L. A. Hawkes, A. Fahlmann, Katsufumi Sato. Introduction to the theme issue: measuring physiology in free-living animals. Philosophical Transaction Royal Society B 376: 20200210 (2021).
  •         L. A. Hawkes, A. Fahlmann, Katsufumi Sato. Why is physiologging? Introduction to the theme issue, part 2. Philosophical Transaction Royal Society B 376: 20200228 (2021).
  •         J. Chris McKnight, Eric Mulder, Alexander Ruesch, Jana M. Kainerstorfer, Jingyi Wu, Naser Hakimi, Steve Balfour, Mathijs Bronkhorst, Jorn M. Horschig, Frank Pernett, Katsufumi Sato, Gordon D. Hastie, Peter Tyack, Erika Schagatay. When the human brain goes diving: using near-infrared spectoroscopy to measure cerebral and systemic cardiovascular responses to deep, breath-hold diving in elite freedivers. Philosophical Transaction Royal Society B 376: 20200349 (2021).
  •         Hiromi Kudo, Minami W Okuyama, Kentaro Q Sakamoto, Kei Uchida, Katsufumi Sato. Serotonin-manipulated juvenile green sea turtles (Chelonia mydas) exhibit reduced fear-like behaviour. Endangered Species Research 46: 185-191 (2021).
  •         Aran Garrod, Sei Yamamoto, Kentaro Q. Sakamoto, Katsufumi Sato. Video and acceleration records of streaked shearwaters allows detection of two foraging behaviours associated with large marine predators. PLoS ONE 16: e0254454 (2021).
  •         Shigenori Nobata, Takashi Kitagawa, Shouji Houki, Motohiro Ito, Yoshinori Aoki, Katsufumi Sato, Susumu Hyodo. Rrelationships between maturational status and migration behavior of homing chum salmon Oncorhynchus keta in inner bays of the Sanriku Coast. General and Comparative Endocrinology 313 (2021).


2020

  •         Tomoko Hamabata, Ayumi Matsuo, Mitsuhiko P. Sato, Satomi Kondo, Kazunari Kameda, Isao Kawazu, Takuya Fukuoka, Katsufumi Sato, Yoshihisa Suyama, Masakado Kawata. Natal origin identification of green turtles in the North Pacific by genome-wide population analysis with limited DNA samples. Frontiers in Marine Science (2020).
  •         G. Kooyman, K. Goetz, C. L. Willams, P. J. Ponganis, K. Sato, S. Eckert, M. Horning, P. T. Thorson, R. P. van Dam. Crary bank: a deep foraging habitat for emperor penguins in the western Ross Sea. Polar Biology 43: 801-811 (2020).
  •         Andreas Fahlman, Katsufumi Sato, and Patrick Miller. Improving estimates of diving lung volume in air-breathing marine vertebrates.Journal of Experimental Biology 223: jeb216846 (2020).
  •         Itsumi Nakamura, Rui Matsumoto, and Katsufumi Sato. Body temperature stability observed in the whale sharks, the world’s largest fish. Journal of Experimental Biology 223: jeb210286 (2020).
  •         Yoshida et al. Underwater video and still-image dataset of fishes and other aquatic animals in Lake Biwa, Japan, observed via carp-mounted video loggers. Ecological Research (in press).
  •         Shohei Kobayashi, Takuya Fukuoka, Chihiro Kinoshita, Katsufumi Sato, Katsuhiko Arai and Gen Watanabe. Male-biased Sex Ratio of Adult-sized Loggerhead Turtles (Caretta caretta) Migrating to High-latitude Summer-restricted Foraging Grounds in the Northwest Pacific.Pacific Science 74: 1-13 (2020).

2019


        Takeshi Doi, Andrea Storto, Takuya Fukuoka, Hiroshi Suganuma and Katsufumi Sato. Impacts of temperature measurements from sea turtles on seasonal prediction around the Arafura Sea. Frontiers in Marine Science 6:719 (2019).
        Takuya Fukuoka, Chihiro Kinoshita, Katsufumi Sato. Northernmost bycatch record of an olive ridley turtle (Lepidochelys olivacea) in the Pacific coast of Japan. Marine Turtle Newsletter 159: 17-22 (2019).
        Ricardo Domingues, Akira Kuwano-Yoshida, Patricia Chardon-Maldonado, Robert E. Todd, George Halliwell, Hyun-Sook Kim, I.-I. Lin, Katsufumi Sato, Tomoko Narazaki, Lynn K. Shay, Travis Miles, Scott Glenn, Jun A. Zhang, Steven R. Jayne, Luca Centurioni, Matthieu Le Hénaff, Gregory R. Foltz, Francis Bringas, M. M. Ali, Steven F. DiMarco, Shigeki Hosoda, Takuya Fukuoka, Benjamin LaCour, Acichal Mehra, Elizabeth R. Sanabia, John R. Gyakun, Jili Dong, John A. Knaff and Gustavo Goni. Ocean observations in support of studies and forecasts of tropical and extratropical cyclones. Frontiers in Marine Science 6: 446 (2019).
        Nobata et al. Spreading of river water guides migratory behavior of homing chum salmon Oncorhynchus keta in Otsuchi Bay, a narrow inlet with multiple river flows. Zoological Science36: 449-457 (2019).
        Rob Harcourt et al., Animal-Borne Telemetry: an integral component of the ocean observing toolkit. Frontiers in Marine Science 6:326.
       Takuya Fukuoka, Kazuyoshi Omuta, Toshio Hidaka, Katsufumi Sato. First record of a known-age juvenile loggerhead turtle (Caretta caretta) at the Sanriku Coast in the Northwest Pacific Ocean. Coastal Marine Science 42: 8-11.
       Kozue Shiomi, Katsufumi Sato, Nobuhiro Katsumata and Ken Yoda. Temporal and spatial determinants of route selection in homing seabirds. Behaviour. 156: 1165-1183.
       Cassondra L. Williams, Katsufumi Sato and Paul J. Ponganis. Activity not submergence explains diving heart rates of captive loggerhead turtles. Journal of Experimental Biology 222: jeb200824.
       Takaaki K. Abe, Takashi Kitagawa, Yuya Makiguchi, Katsufumi Sato. Chum salmon migrating upriver accommodate to environmental temperatures through metabolic compensation. Journal of Experimental Biology
        Yu Akiyama, Tomonari Akamatsu, Marianne H. Rasmussen, Maria R. Iversen, Takashi Iwata, Yusuke Goto, Kagari Aoki, Katsufumi Sato. Leave or stay? Video-loggers revealed foraging efficiency of humpback whales under temporal change in prey density. PLoS One
       Takuya Fukuoka, Tomoko Narazaki, Chihiro Kinoshita and Katsufumi Sato. Diverse foraging habits of juvenile green turtles (Chelonia mydas) in a summer-restricted foraging habitat in the northwest Pacific Ocean. Marine Biology


2018

       Yasumasa Miyazawa, Akira Kuwano-Yoshida, Takeshi Doi, Hatsumi Nishikawa, Tomoko Narazaki, Takuya Fukuoka, Katsufumi Sato. Temperature profiling measurements by sea turtles improve ocean state estimation in the Kuroshio-Oyashio confluence region. Ocean Dynamics (in press).
       Rei Yamashita, Hideshige Takada, Arisa Nakazawa, Akinori Takahashi, Motohiro Ito, Takashi Yamamoto, Yuuki Watanabe, Nobuo Kokubun, Katsufumi Sato, Sarah Wanless, Francis Daunt, David Hyrenbach, Michelle Hester, Tomohiro Deguchi, Bungo Nishizawa, Akiko Shoji, Yutaka Watanuki. Global monitoring of persistent organic pollutants (POPs) using seabird preen gland oil. Archives of Environmental Contamination and Toxicology 75: 545-556 (2018).
       Miho Sakao, Hirohiko Takeshima, Koji Inoue, Katsufumi Sato. Extra-pair paternity in socially monogamous Streaked Shearwater: forced copulation or female solicitation? Journal of Ornithology.
        Tomoko Narazaki, Saana Isojunno, Douglas P Nowacek, Rene Swift, Ari S Friedlaender, Christian Ramp, Sophie Smout, Kagari Aoki, Volker B Deecke, Katsufumi Sato, Patrick JO Miller. Body density of humpback whales (Megaptera novaeangliae) in feeding aggregations estimated from hydrodynamic gliding performance. PLoS One (in press).
        Chihiro Kinoshita, Takuya Fukuoka, Yasuaki Niizuma, Tomoko Narazaki and Katsufumi Sato. High resting metabolic rates with low thermal dependence induce active dives in overwintering Pacific juvenile loggerhead turtles. Journal of Experimental Biology (in press).
        Olivia Hicks, Sarah Burthe, Francis Daunt, Mark Newell, Adam Butler, Motohiro Ito, Katsufumi Sato, Jonathan A. Green. The energetic cost of parasitism in a wild population. Proceeding of Royal Society B 285: 20180489.(2018).
        Nicholas L. Payne, Carl G. Meyer, James A. Smith, Jonathan D. R. Houghton, Adam Barnett, Bonnie J. Holmes, Itsumi Nakamura, Yannis P. Papastamatiou, Mark A. Royer, Daniel M. Coffy, James M. Anderson, Melanie R. Hutchinson, Katsufumi Sato, Lewis G. Halsey. Combining abundance and performance data reveals how temperature regulates coastal occurrences and activity of a roaming apex predator. Global Change Biology 2018: 1-10 (2018).


2017

        Takashi Iwata, Tomonari Akamatsu, Surasak Thongsukdee, Phaothep Cherdsukjai, Kanjana Adulyanukosol, Katsufumi Sato. Tread-water feeding of Bryde’s whales. Current Biology 27: R1141-R1155 (2017)
        Yusuke Goto, Ken Yoda, Katsufumi Sato. Asymmetry hidden in birds’ tracks reveals wind, heading, and orientation ability over the ocean. Science Advances 3, e1700097 (2017). DOI: 10.1126/sciadv.1700097
       Kagari Aoki, Katsufumi Sato, Saana Isojunno, Tomoko Narazaki, Patrick J. O. Miller. High diving metabolic rate indicated by high-speed transits to depth in negatively buoyant long-finned pilot whales. Journal of Experimental Biology 220: 3802-3811 (2017). DOI: 10.1242/jeb.158287
       Makoto A. Yoshida, Daisuke Yamamoto, Katsufumi Sato. Physostomous channel catfish, Ictalurus punctatus, modify swimming mode and buoyancy according to flow conditions. Journal of Experimental Biology 220: 597-606 (2017)


2016

       Kozue Shiomi, Katsufumi Sato, Yves Handrich, Charles A. Bost. Diel shift of swim speeds in king penguins in relation to changes of light intensity. MEPS 561: 233-243 (2016). DOI: 10.3354/meps11930
       Yoshinari Yonehara, Yusuke Goto, Ken Yoda, Yutaka Watanuki, Lindsay C. Young, Henri Weimerskirch, Charles-André Bost, Katsufumi Sato. Flight paths of seabirds soaring over the ocean surface enable measurement of fine-scale wind speed and direction. PNAS (2016). DOI: 10.1073/pnas.1523853113.
       Patrick J. O. Miller, Tomoko Narazaki, Saana Isojunno, Kagari Aoki, Sophie Smout, and Katsufumi Sato. Body density and diving gas volume of the northern bottlenose whale (Hyperoodon ampullatus). Journal of Experimental Biology 219: 2458-2468 (2016)
       Takuya Fukuoka, Misaki Yamane, Chihiro Kinoshita, Tomoko Narazaki, Greg J. Marshall, Kyler J. Abernathy, Nobuyuki Miyazaki, Katsufumi Sato. The feeding habit of sea turtles influences their reaction to artificial marine debris. Scientific Reports 6: 28015 (2016).
       Naoyuki Miyata, Tomohiko Mori, Masaaki Kagehira, Nobuyuki Miyazaki, Michihiko Suzuki and Katsufumi Sato. In situ CTD measurement using fish-borne recorders reveal short-term excursions of Japanese sea bass from sea water to freshwater. Aquatic Biology 219: 2458-2468 (2016).
        Anthony A. Robson, Lewis G. Halsey and Laurent Chauvaud. Feet, heat and scallops: what is the cost of anthropogenic disturbance in bivalve aquaculture? Royal Society Open Science 3: 150679 (2016)
       Takashi Kitagawa, Susumu Hyodo and Katsufumi Sato. Atmospheric depression-mediated water temperature changes affect the vertical movement of chum salmon Oncorhynchus keta. Marine Environmental Research 119: 72-78 (2016)
        Daisuke Ochi, Kei Matsumoto, Nariko Oka, Tomohiro Deguchi, Katsufumi Sato, Takashi P. Satoh, Fumihito Muto, Yutaka Watanuki. Dual foraging strategy and chick growth of streaked shearwater Calonectris leucomelas at the two colonies of different oceanographic environment. Ornithological Science (in press)
       Manfred R. Enstipp, Katia Ballorain, Stéphane Ciccione, Tomoko Narazaki, Katsufumi Sato, Jean-Yves Georges, Energy expenditure of adult green turtles (Chelonia mydas) at their foraging grounds and during simulated oceanic migration. Functional Ecology (2016)
        Graeme C. Hays, Luciana C. Ferreira, Ana M.M. Sequeira, Mark G. Meekan, Carlos M. Duarte, Helen Bailey, Fred Bailleul, W. Don Bowen, M. Julian Caley, Daniel P. Costa, Victor M. Eguíluz, Sabrina Fossette, Ari S. Friedlaender, Nick Gales, Adrian C. Gleiss, John Gunn, Rob Harcourt, Elliott L. Hazen, Michael R. Heithaus, Michelle Heupel, Kim Holland, Markus Horning, Ian Jonsen, Gerald L. Kooyman, Christopher G. Lowe, Peter T. Madsen, Helene Marsh, Richard A. Phillips, David Righton, Yan Ropert-Coudert, Katsufumi Sato, Scott A. Shaffer, Colin A. Simpfendorfer, David W. Sims, Gregory Skomal, Akinori Takahashi, Philip N. Trathan, Martin Wikelski, Jamie N. Womble and Michele Thums. Key questions in marine megafauna movement ecology. TREE 31: 463-475 (2016).
       Yasumasa Miyazawa, Xinyu Guo, Sergey M. Varlamov, Toru Miyama, Ken Yoda, Katsufumi Sato and Keiji Sato. Assimilation of the seabird and ship drift data in the north-eastern sea of Japan into an operational ocean nowcast/forecast system. Scientific Reports5: 17672 (2015).
       Yukihisa Kogure, Katsufumi Sato, Yutaka Watanuki, Sarah Wanless and Francis Daunt. European shags optimize their flight behavior according to wind conditions. Journal of Experimental Biology 219:311-318 (2016).


2015

       Aurélie Jolivet, Laurent Chauvaud, Julien Thébault, Anthony A. Robson, Pascal Dumas, George Amos and Anne Lorrain. Circadian behavior of Tectus (Trochus) niloticus in the southwest Pacific inferred from accelerometry. Movement Ecology 3:26 (2015)
       Tomohiko Mori, Naoyuki Miyata, Jun Aoyama, Yasuaki Niizuma and Katsufumi Sato. Estimation of metabolic rate from activity measured by recordes deployed on Japanese sea bass Lateolabrax japonicus. Fisheries Science 81:871-882 (2015).
       Iwata. T., Sakamoto. K. Q., Edwards. E. W. J., Staniland. I. J., Trathan. P. N., Goto. Y., Sato. K., Naito. Y., Takahashi. A. The influence of preceding dive cycles on the foraging decisions of Antarctic fur seals. Biology Letters 11: 20150227 (2015).
        Itsumi Nakamura, Carl Meyer and Katsufumi Sato. Unexpected positive buoyancy in deep sea sharks, Hexanchus griseus, and a Echinorhinus cookei. PLoS One e0127667 (2015)
       Tomoko Narazaki, Katsufumi Sato and Nobuyuki Miyazaki. Summer migration to temperate foraging habitats and active winter diving of juvenile loggerhead turtles Caretta caretta in the western North Pacific. Marine Biology 162:1251-1263(2015)
       Kagari Aoki, Masao Amano, Tsunemi Kubodera, Kyoichi Mori, Ryosuke Okamoto, Katsufumi Sato. Visual and behavioral evidence indicates active hunting by sperm whales. Marine Ecology Progress Series 523:233-241 (2015)
       Itsumi Nakamura, Yusuke Goto, Katsufumi Sato. Ocean sunfish rewarm at the surface after deep excursions to forage for siphonophores. Journal of Animal Ecology 84:590-603 (2015)
       Yu Akiyama, Yoshio Matsuda, Natsuko Sakurai, Katsufumi Sato. Evaluation of wave drag on bottlenose dolphin Tursiops truncatus from swimming effort. Coastal Marine Science in press (2015)
       Otaki Takayoshi, Masahiro Hamana, Hideaki Tanoue, Nobuyuki Miyazaki, Takuro Shibuno and Teruhisa Komatsu. Three-dimensional mapping of red stingray (Dasyatis akajei) movement with reference to bottom topography. Ocean Science Journal in press (2015)
       Takuya Fukuoka, Tomoko Narazaki, Katsufumi Sato. Summer-restricted migration of green turtles (Chelonia mydas) to a temperate habitat of the northwest Pacific Ocean. Endangered Species Research 28:1-10 (2015)




2014


       Christopher D. Marshall, John Wang, Axayacatl Rocha-Olivares, Carlos Godines-Reyes, Shara Fisler, Tomoko Narazaki, Katsufumi Sato and Blair D. Sterba-Boatwright. Scaling of bite performance with head and carapace morphometrics in green turtles (Chelonia mydas). Journal of Experimental Marine Biology and Ecology 451:91-97 (2014)
       Takashi Yamamoto, Akinori Takahashi, Katsufumi Sato, Nariko Oka, Maki Yamamoto and Philip N. Trathan. Individual consistency in migratory behaviour of a pelagic seabird. Behaviour 151: 683-701(2014).
       Ken Yoda, Kozue Shiomi and Katsufumi Sato. Foraging spots of streaked shearwaters in relation to ocean surface currents as identified using their drift movements. Progress in Oceanography 122: 54-64(2014)
       Maria I. Bogdanova, Sarah Wanless, Michael P. Harris, Ian Lindstrom, Adam Butler, Mark A. Newell, Katsufumi Sato, Yutaka Watanuki, Matt Parsons and Francis Daunt. Among-year and within-population variation in foraging distribution of European shags Phalacrocorax aristotelis over two decades: implications for marine spatial planning. Biological Conservation 170: 292-299 (2014)
       Hideaki Nishizawa, Tomoko Narazaki, Takuya Fukuoka, Katsufumi Sato, Tomoko Hamabata, Masato Kinoshita, Nobuaki Arai. Genetic composition of loggerhead turtle feeding aggregations: Insight into migration patterns in the North Pacific. Endangered Species Research 24: 85-93 (2014)
       Hideaki Nishizawa, Tomoko Narazaki, Takuya Fukuoka, Katsufumi Sato, Masato Kinoshita and Nobuaki Arai. Juvenile green turtles in the northern edge: mtDNA evidence of long-distance westward dispersals in the Northern Pacific Ocean. Endangered Species Research 24: 171-179 (2014)
       Masao Amano, Aya Kourogi, Kagari Aoki, Motoi Yoshioka and Kyoichi Mori. Differences in sperm whale codas between two waters off Japan: possible geographic separation of vocal clans. Journal of Mammalogy 95: 169-175 (2014)
       Katsufumi Sato. Body temperature stability achieved by the large body mass of sea turtles. Journal of Experimental Biology 217: 3607-3614 (2014)
       Ippei Suzuki, Katsufumi Sato, Andreas Fahlman, Yasuhiko Naito, Nobuyuki Miyazaki and Andrew W. Trites. Drag, but not buoyancy, affects swim speed in captive Steller sea lions. Biology Open 3: 379-386 (2014)
       Itsumi Nakamura and Katsufumi Sato. Ontogenetic shift in foraging habit of ocean sunfish Mola mola from dietary and behavioral studies. Marine Biology 161: 1263-1273 (2014)
       Fleur Visser, Patrick J.O. Miller, Ricardo N. Antunes, Machiel G. Oudejans, Monique L. Mackenzie, Kagari Aoki, Frans-Peter A. Lam, Petter H. Kvadsheim, Jef Huisman and Peter L. Tyack. The social context of individual foraging behaviour in long-finned pilot whales (Globicephala melas). Behaviour 151: 1453–1477 (2014).


2013


       Atsuo Ito, Rei Yamashita, Hideshige Takada, Takashi Yamamoto, Kozue Shiomi, Carlos Zavalaga, Tomoya Abe, Shinichi Watanabe, Maki Yamamoto, Katsufumi Sato, Hiroyoshi Kohno, Tomohiko Iida, Ken Yoda and Yutaka Watanuki. Contaminants in tracked seabirds show regional patterns of marine pollution. Environmental Science & Technology 47: 7862-7867 (2013).
       Tomoko Narazaki, Katsufumi Sato, Kyler J. Abernathy, Greg J. Marshall and Nobuyuki Miyazaki. Loggerhead turtles (Caretta caretta) use vision to forage on gelatinous prey in mid-water. PLoS ONE 8: e66043 (2013). Doi: 10.1371/journal.pone.0066043.
       Katsufumi Sato, Kagari Aoki, Yuuki Y. Watanabe and Patrick J. O. Miller. Neutral buoyancy is optimal to minimize the cost of transport in horizontally swimming seals. Scientific Reports 3:2205; DOI:10.1038/srep02205 (2013).
       Kagari Aoki, Mai Sakai, Patrick J. O. Miller, F. Visser and Katsufumi Sato. Body contact and synchronous diving in long-finned pilot whales. Behavioural Processes 99:12-20 (2013).
       Takashi Iwata, Shiroh Yonezaki, Kaoru Kohyama and Yoko Mitani. Detection of grooming behaviours with an acceleration data logger in a captive northern fur seal (Callorhinus ursinus). Aquatic Mammals (2013).
       Kentaro Q. Sakamoto, Akinori Takahashi, Takashi Iwata, Takashi Yamamoto, Maki Yamamoto and Philip N. Trathan. Heart rate and estimated energy expenditure of flapping and gliding in black-browed albatrosses. The Journal of Experimental Bilogy. (2013)
       M Amano, A Kourogi, K Aoki, M Yoshioka, M Kyoichi. Differences in sperm whale codas between two waters off Japan: possible geographic separation of vocal clans. Journal of Mammalogy, in press.


2012


       Ken YODA, Tadashi Tajima, Sachiho Sasaki, Katsufumi Sato and Yasuaki Niizuma. Influence of Local Wind Conditions on the Flight Speed of the Great Cormorant Phalacrocorax carbo. International Journal of Zoology Doi: 10.1155/2012/187102 (2012).
       Shinichi Watanabe, Katsufumi Sato and Paul J. Ponganis. Activity time budget during foraging trips of emperor penguins. PLoS ONE 7: e50357. Doi: 10.1371/journal.pone.0050357 (2012).
       Ichiro Ken Shimatani, Ken Yoda, Nobuhiro Katsumata and Katsufumi Sato. Toward the quantification of a conceptual framework for movement ecology using circular statistical modeling. PLoS ONE 7: e50309. Doi:10.1371/journal.pone.0050309 (2012).
       Christopher D Marshall, Alejandra Guzman, Tomoko Narazaki, Katsufumi Sato, Emily A. Kane, and Blair Sterba-Boatwright. The ontogenetic scaling of bite force and head size in loggerhead sea turtles (Caretta caretta): implications for durophagy in neritic, benthic habitats. Journal of Experimental Biology 215: 4166-4174 (2012).
       Ryota Hayashi, Benny K. K. Chan, Noa Simon-Blecher, Hiromi Watanabe, Tamar Guy-Haim, Takahiro Yonezawa, Yaniv Levy, Takuho Shuto and Yair Achituv. Phylogenetic position and evolutionary history of the turtle and whale barnacles (Cirripedia: Balanomorpha: Coronuloidea). Molecular Phylogenetics and Evolution 67: 9-14 (2013).
       Yoshio Takei, Hiroshi Miyanishi, Shigenori Nobata, Marty K. S. Wong, Taro Watanabe, Albert Ventura, Aya Shiozawa, Susumu Hyodo and Makoto Kusakabe. Establishment and validation of an aquarium system to evaluate salinity preference in conscious rainbow trout. Coastal Marine Science (in press).


2011


       Chan BKK and Hayashi R. Epibiotic barnacles (Crustacea: Cirripedia) collected in the Kumejima 2009 expedition, with descriptions of two new species. Zootaxa (in press).
       Shuto T and Hayashi R. Floating castles in offshore waters: the barnacles of two giant Fish Aggregation Devices from off Okinawa, Japan. Landsc Ecol Eng DOI: 10.1007/s11355-012-0190-7 (in press).
       Yamamoto T, Takahashi A, Oka N, Shirai M, Yamamoto M, Katsumata N, Sato K, Watanabe S and Trathan PN. Inter-colony differences in the incubation pattern of streaked shearwaters in relation to the local marine environment. Waterbirds (in press).
       Williams CL, Sato K, Shiomi K and Ponganis PJ. Muscle energy stores and stroke rates of emperor penguins: Implications for muscle metabolism and dive performance. Physiol Biochem Zool (in press).
       Aoki K, Amano M, Mori K, Kourogi A, Kubodera T and Miyazaki N. Active hunting by deep-diving sperm whales: 3D dive profiles and maneuvers during bursts of speed. Mar Ecol-Prog Ser 444:289-301 (2012).
       Hayashi R. Atlas of the barnacles on marine vertebrates in Japanese waters including taxonomical review of superfamily Coronuloidea (Cirripedia: Thoracica). J Mar Biol Assoc UK 92: 107-127 (2012).
       Shiomi K, Sato K and Ponganis PJ. Point of no return in diving emperor penguins: is decision time to return limited by the number of strokes? J Exp Biol 215: 135-140 (2012).
       Shiomi K, Yoda K, Katsumata N and Sato K. Temporal tuning of homeward flight in seabirds. Anim Behav 83: 355-359 (2012).
       Tanoue H, Komatsu T, Tsujino T, Suzuki I, Watanabe M, Goto H and Miyazaki N. Feeding events of Japanese lates Lates japonicus detected by a high-speed video camera and three-axis micro-acceleration data-logger, Fisheries Sci (on line first published)(2012).
       Alabsi NM, Tanoue H, Komatsu T, Charef A, Mitani I, Kato M, Horii T, Aoki I, Miyazaki N. Measurement of the swimming behavior of a deep-water fish, the Splendid Alfonsino (Beryx splendens), in captivity using micro data loggers, J Fisheries Aquat Sci 6: 309-321 (2011).
       Aoki K, Watanabe YY, Crocker DE, Robinson PW, Biuw M, Costa DP, Miyazaki N, Fedak MA and Miller PJO. Northern elephant seals adjust gliding and stroking patterns with changes in buoyancy: validation of at-sea metrics of body density. J Exp Biol 214: 2973-2987 (2011).
       Hayashi R, Narazaki T, Takuma S and Sato K. Chelonia mydas agassizii (Black [Pacific] Seaturtle). Epibiont barnacles. Herpetol Rev 42: 264-265 (2011).
       Inoue K, Miyanishi H, Nobata S and Takei Y. Evolutionary implication of the absence of atrial natriuretic peptide (ANP) in euryhaline Oryzias fishes. Environ Biol Fish DOI: 10.1007/ s10641-011-9852-9 (2011).
       Kikuchi M, Vera M. F. de Silva, Rosas FCW, Souza D and Miyazaki N. The implications of turning behaviour performed by Amazonian manatees after release into the wild. J Ethol 30: 187-190 (2011).
       Miyanishi H, Nobata S and Takei Y. Relative antidipsogenic potencies of six homologous natriuretic peptides in eels. Zool Sci 28: 719-726 (2011).
       Nakamura I, Watanabe YY, Papastamatiou YP, Sato K and Meyer CG. Yo-yo vertical movements suggest a foraging strategy for tiger sharks Galeocerdo cuvier. Mar Ecol-Prog Ser 424: 237-246 (2011).
       Nobata S, Donald JA, Balment RJ and Takei Y. Potent cardiovascular effects of homologous urotensin II (UII)-related peptide and UII in unanesthetized eels after peripheral and central injections. Am J Physiol Regul Integr Comp Physiol 300: R437-446 (2011).
       Nobata S and Takei Y. The area postrema is a central player for regulation of drinking behavior in Japanese eels. Am J Physiol Regul Integr Comp Physiol 300: R1569-1577 (2011).
       Sakai M, Aoki K, Sato K, Amano M, Baird RW, Webster DL, Schorr GS and Miyazaki N. Swim speed and acceleration measurements of short-finned pilot whales (Globicephala macrorhynchus) in Hawai‘i. Mammal Study 36: 55-59 (2011).
       Sakai M, Wang D, Wang K, Li S and Akamatsu T. Do porpoises choose their associates? A new method for analyzing social relationships among cetaceans. PLoS ONE 6(12): e28836. doi:10.1371/journal.pone.0028836 (2011).
       Samarra FP, Fennell A, Aoki K, Deecke VB and Miller PJO. Persistence of skin marks on killer whales (Orcinus orca) caused by the parasitic sea lamprey (Petromyzon marinus) in Iceland. Mar mamm sci Article first published online: DOI: 10.1111/j.1748-7692.2011.00486.x (2011).
       Sato K, Shiomi K, Marshall G, Kooyman GL and Ponganis PJ. Stroke rates and diving air volumes of emperor penguins: implications for dive performance. J Exp Biol 214: 2854-2863 (2011).
       Yamamoto T, Takahashi A, Oka N, Iida T, Katsumata N, Sato K and Trathan PN. Foraging areas of streaked shearwaters in relation to seasonal changes in the marine environment of the Northwestern Pacific: inter-colony and sexual differences. Mar Ecol-Prog Ser 424: 191-204 (2011).
       Yuji K. Fuse, Kenaro Q. Sakamoto, Katsufumi Sato and Yoshiaki Habara. Cardiorespiratory pattern of rest-associated apnea in a Weddell seal: a case study at an ice hole in Antarctica. Polar Biol DOI 10.1007/s00300-011-1140-9 (2011).
       Watanabe KK, Hatase H, Kinoshita M, Omuta K, Bando T, Kamezaki N, Sato K, Matsuzawa Y, Goto K, Nakashima Y, Takeshita H, Aoyama J and Tsukamoto K. Population structure of the loggerhead turtle Caretta caretta, a large marine carnivore that exhibits alternative foraging behaviors. Mar Ecol-Prog Ser 424: 273-283 (2011).
       Watanabe YY, Takahashi A, Sato K, Viviant M and Bost C-A. Poor flight performance in deep-diving cormorants. J Exp Biol 214: 412-421 (2011).


2010


       Katsufumi Sato, Kozue Shiomi, Greg Marshall, Gerald L. Kooyman and Paul J. Ponganis. (2011) Stroke rates and diving air volumes of emperor penguins: implications for dive performance. Journal of Experimental Biology
       Natheer Mohammad Alabsi, Hideaki Tanoue, Teruhisa Komatsu, Aymen Charef, Isamu Mitani, Mitsuhiro Kato, Toyomitsu Horii , Ichiro Aoki and Nobuyuki Miyazaki. (2011) Measurement of the swimming behavior of a deep-water fish, the splendid alfonsino (Beryx splendens), in captivity using micro data loggers. Journal of Fisheries and Aquatic Science. 6: 309-321.
       Takashi Yamamoto, Akinori Takahashi, Nariko Oka, Takahiro Iida, Nobuhiro Katsumata, Katsufumi Sato and Phillip N. Trathan. (2011) Foraging areas of streaked shearwaters in relation to seasonal changes in the marine environment of the Northwestern Pacific: inter-colony and sexual differences. Marine Ecology Progress Series. 424: 191-204.
       Kunihiro K. Watanabe, Hideo Hatase, Masato Kinoshita, Kazuyoshi Omuta, Takeharu Bando, Naoki Kamezaki, Katsufumi Sato, Yoshimasa Matsuzawa, Kiyoshi Goto, Yoshito Nakashima, Hiroshi Takeshita, Jun Aoyama and Katsumi Tsukamoto. (2011) Population structure of the loggerhead turtle Caretta caretta, a large marine carnivore that exhibits alternative foraging behaviors. Marine Ecology Progress Series. 424: 273-283.
       Moe Matsumura, Yuuki Y. Watanabe, Patrick W. Robinson, Patrick J. O. Miller, Daniel P. Costa and Nobuyuki Miyazaki. (2011) Underwater and surface behavior of homing juvenile northern elephant seals. Journal of Experimental Biology. 214: 629-636.
       Takashi Yamamoto, Akinori Takahashi, Nobuhiro Katsumata, Katsufumi Sato and Phillip N. Trathan. (2010) At-sea distribution and behavior of streaked shearwaters (Calonectris leucomelas) during non-breeding period: from temperate to tropical oceans. Auk. 127: 871-881.
       Watanabe YY, Takahashi A, Sato K, Viviant M and Bost C-A. Poor flight performance in deep-diving cormorants. Journal of Experimental Biology. (2011) 214: 412-421.
       Mai Sakai, Kagari Aoki, Katsufumi Sato, Masao Amano, Robin W. Baird, Daniel L. Webster, Gregory S. Schorr, and Nobuyuki Miyazaki. (2011) Swim speed and acceleration measurements of short-finned pilot whales (Globicephala macrorhynchus) in Hawai‘i. Mammal Study. 36: 55-59
       Y. Y. Watanabe, K. Sato, Y. Watanuki, A. Takahashi, Y. Mitani, M. Amano, K. Aoki, T. Narazaki, T. Iwata, S. Minamikawa, N. Miyazaki. (2011) Scaling of swim speed in breath-hold divers. Journal of Animal Ecology. 80: 57-68.
       Mumi Kikuchi, Kentaro Q. Sakamoto, Katsufumi Sato. (2010) Acquisition of gliding skills by Weddell seal (Leptonychotes weddellii) pups during lactation. Polar Biology. 33: 1429-1435.
       Kozue Shiomi, Tomoko Narazaki, Katsufumi Sato, Kenichiro Shimatani, Nobuaki Arai, Paul J. Ponganis and Nobuyuki Miyazaki. (2010) Data-processing artefacts in 3-D dive path reconstruction from geomagnetic and acceleration data. Aquatic Biology. 8: 299-304
       Morisaka, T., Kohshima, S., Yoshioka, M., Suzuki, M., & Nakahara, F. (2010) Recent studies on captive cetaceans in Japan: Working in tandem with studies on cetaceans in the wild. International Journal of Comparative Psychology. 23: 644-663.
       Sakamoto K, Sato K, Kato A, Fukui D, Bando G, Naito Y, Habara Y, Ishizuka M and Fujita S. (2010)Metabolic alkalosis due to feeding chicks in breeding Adélie penguins Pygoscelis adeliae under natural conditions. Physiological and Biochemical Zoology. 83: 232-238.
       Mumi Kikuchi; Vera M. F. da Silva; Fernando C. W. Rosas; Nobuyuki Miyazaki. (2010) Application of acceleration data loggers to classify the behavior of captive Amazonian manatees (Trichechus inunguis). Coastal marine science. 34: 24-30


2009


       Sakamoto KQ, Sato K, Ishizuka M, Watanuki Y, Takahashi A, Daunt F, Wanless S. (2009) Can Ethograms Be Automatically Generated Using Body Acceleration Data from Free-Ranging Birds? PLos One, 4: e5379.
       Sato K, Sakamoto KQ, Watanuki Y, Takahashi A, Katsumata N, Bost C-A, Weimerskirch H. (2009) Scaling of Soaring Seabirds and Implications for Flight Abilities of Giant Pterosaurs. PLoS ONE 4: e5400.
       Suzuki I, Naito Y, Folkow LP, Miyazaki N, Arnoldus SB. (2009) Validation of a device for accurate timing of feeding events in marine animals, Polar Biology, 32: 667-671.
       Watanabe Y, Lydersen C, Sato, Naito Y, Miyazaki N, Kovacs KM. (2009) Diving behavior and swimming style of nursng bearded seal pups, Marine ecology progress series, 380: 287-294.
       Weimerskirch H, Shaffer SA, Tremblay Y, Costa DP, Gadenne H, Kato A, Ropert-coudert Y, Sato K, Aurioles D. (2009) Species and sex specific differences in foraging behaviour and foraging zones in blue-footed and brown boobies in the Gulf of California. Marine Ecology Progress Series, 391: 267-278
       Sato K, Shiomi K, Watanabe Y, Watanuki Y, Takahashi A, Ponganis PJ. (2009) Scaling of swim sped and stroke frequency in geometrically similar penguins: they swim optimally to minimize cost of transport. Proceedings of the Royal Society B, 274: 471-477.